VMD, Veterinary Medicine, University of Pennsylvania, 1965

MS, Dairy Science, The Pennsylvania State University, 1969

BS, Dairy Science, The Pennsylvania State University, 1961

General animal health, herd health management, cardiac and respiratory physiology

Courses Taught

V. Sc. 050S Mechanisms of Disease

V. Sc. 303 Principles of Animal Disease

V. Sc. 303H Principles of Animal disease Honors

V. Sc. 407 Dairy Herd Health Programs

Bovine reproductive physiology, reproductive diseases

Relationships Between Patterns of Follicular Development and Fertility. NE-161 Regional Project

Effect of Uterine Infection on Patterns of Ovarian Follicular Development and Sunsequent Fertility. Penna. Department of Agriculture.

Smith, RM, RM Leach, LD Muller, LC Griel Jr, DE Baker. Effects of long-term dietary cadmium chloride on tissue, milk, and urine mineral concentrations of dairy cattle. J Animal Sci 69:4088-4096, 1991.

Jerry, DL, LC Griel Jr, JF Kavanaugh, RS Kensinger. Binding and bioactivity of ovine and porcine prolactins in porcine mammary tissue. J Endocrinology 130:43-51, 1991.

Henault, MA, GJ Killian, JF Kavanaugh, LC Griel Jr. Effect of accessory sex gland fluid from bulls of differing fertilities on the ability of cauda epididymal sperm to penetrate zona-free bovine oocytes. Biol Reprod 52:390-397, 1995.

Cooperative Regional Research Project NE-161. Relationship of Fertility to Patterns of Ovarian Follicular Development and Associated Hormone Profiles in Dairy Cows and Heifers. J. Animal Sci. 74: 1943-1952, 1996.

Way, Amy L., L. C. Griel Jr. and G. J. Killian. Effects of Accessory Sex Gland Fluid on Viability, Capacitation and the Acrosome Reaction of Cauda Epididymal Bull Spermatozoa. Andrology 21: 213-219, 2000.

Townson, D.H., P.C.W. Tsang, W. R. Butler, M. Frajblat, L. C. Griel Jr., C. J. Johnson, C. M. Niksic and J. C. Pate. J. An. Sci. 80: 1053-1058, 2002